Seasonal Habitat Shifts by Benthic Fishes in Headwater Streams
Fish-habitat associations in streams have been widely studied; however, temporal considerations have been neglected, particularly during the winter. We quantitatively sampled perennial headwater streams in the Missouri Ozarks during the summer (n = 13) and winter (n = 4) to evaluate possible habitat shifts by three benthic fishes at two spatial scales: channel unit and microhabitat. Density of all three headwater species in streams was generally lower in winter than summer, with some species being ubiquitous in channel units of streams during the summer and almost entirely absent from...